Yes, on a Friday afternoon I'm looking for things to procrastinate over, so I decided amuse myself with yet another of these rambles and look at the most troublesome of the Dinobots, Strafe. Why is Strafe troublesome? Remarkably, it's not the two heads. So, working from the POV that Strafe's second head and tail is due to a conjoined abberation...what the hell is he? ------------------------ First of all, Strafe is easily placed in pterosauria based on his being a winged reptile with an elongate rostrum/beak, quadrapedalism, etc etc. I could actually find the paper with all the pterosaur characters but he is definitely a pterosaur. Now things get complicated. He has been announced as being a Pteranodon longiceps, a well known pterosaur from the Niobara Formation in the us. For reference here's the skull of the typical Pteranodon And here's Strafe The skull structure is completely different. He's got teeth, his crest extends quite far in front of his eyes and not very far away from the back of the skull (seen in females and juveniles, but they don't have an anterior crest). His jaw is much shorter jaw full of teeth. If we look at the body, Strafe has very lark foreclaws and an elongate tail. He's nothing like the more lightly built Pteranodon, or any animal in the Pteranodon lineage, with the possible exception of Istiodactylus but that has a highly distinct dentition lacking here. So he's definitely not a Pteranodon. Could he be another "advanced pterosaur". Well no. Even though some Cretaceous pterosaurs retained their teeth, none retained a long tail. So is he something along the line of Rhamphorhynchus or some of the earlier pterosaurs? No again. Strafe has a large "blank area" near the frontal crest which is positioned in the same place as the large skull window seen in "advanced" pterosaurs. This means that Strafe cannot be basal, and in fact shows both basal and derived features. So Strafe is a bizzare hybrid, a chimera of some sort from some unknown group. No. Again. In 2009 we found this animal Darwinopterus is considered a key evolutionary step. It has the head of an [advanced pterosaur and the body of a basal one. It is literally like you chopped the head off of one and glued it onto another. Which is pretty much what Strafe is. So amazingly Strafe is a Darwinopterus type animal, belonging to the family Wukongopteridae. However he is definitely not Darwinopterus. Size apart (Darwinopterus is quite small), he's construction is considerably more robust, with massive forelimbs and a heavy set skull. His jaw is shorter, his teeth do not extend to the tip, his tail is proportionally much longer, he has a unique crest morphology etc. Strafe is the most robust wukongopterid known, and should be put in an entirely new genus and species. Perhaps something like Robustodraco but that's debatable. Regardless, Strafe has the interesting honour of being the first wukongopterid on the big screen. Ever.