"After the Fallen was caught trying to harvest a star from a system with life, he turned on the other Primes and made war on them. In order to prevent the Fallen from harvesting the star, the remaining Primes (including Vector Prime) hid the key to the harvester (Matrix) inside a tomb made of their own bodies and sealed it with their own life essence" Technically he was in Revenge Of The Fallen as one of the many 'Fallen-looking' Primes that appeared