So, it would appear that Nautica's father (and I use that term loosely) is Submarauder. Also, this won't end well for Gridlock and Hightail.
At least two more references, the 1986 movie and Team Fortress 2, and it makes sense with Kup and Blitzwing in context and hindsight.
Something just occurred to me. How does Submarauder keep himself covered in shadows even when lasers are coming directly at him?