It seemed as though that all the decepticons spoke cybertronian with the exception of Barricade and Megatron. As far as the autobots go I assume that they were speaking english when talking amongst each other under the bridge and at Griffith Observatory. However, my wife believes that the scene called for a translation from cybertronian to english. I'm rather curious if anyone else thought it was straight out human language or a translation. I believed that the autobots seemed to be more accepting of the humans so they would possibly speak as they do.